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Very early in his life, Knuckle began carving the Tingle Tower and finished when he was ten years old. He works as a carver in the head of the Tingle Tower. Knuckle is also the creator of the Tingle Statues, small golden statues depicting a stance of Tingle.
When Link visits Outset Island with the Tingle Tuner and travels over to the patch of grass and three stones that are in front of Orca and Sturgeon's house, Knuckle greets him by demanding he finds the outdoor bath, and not to go indoors. Tingle encourages Link to do as his brother said. Knuckle subsequently gives him commands like climbing down the ladder that no one uses, and climbing up the biggest rock on top of the tall hill. To test Link's courage, he asks him to jump off the ledge where the suspension bridge once was. Impressed, he gives Link the Hand-Me-Down Tingle Tuner and appears on Tingle Island.
Knuckle appears in Trilby Highlands, and is part of the Kinstone trading quest. It is revealed that he holds a grudge against David Jr., a "Tingle Brother" who is not biologically related to himself, Tingle and Ankle.
In Hyrule Warriors Legends, Tingle's Standard Outfit (Great Sea) is a Knuckle Recolor which can be obtained on the Great Sea Adventure Map.
| Ezlo · King Daltus · Link · Princess Zelda · Smith · Vaati|
—Full list of characters
|Big Green ChuChu · Gleerok · Mazaal · Big Octorok · Gyorg Pair · Vaati Reborn · Vaati Transfigured · Vaati's Wrath|
| Castor Wilds · Cloud Tops · Eastern Hills · Hyrule Town · Lake Hylia · Lon Lon Ranch · Minish Woods · Mount Crenel · Mt. Crenel's Base · North Hyrule Field · Royal Valley · South Hyrule Field · Trilby Highlands · Veil Falls · Western Wood · Wind Ruins|
|Deepwood Shrine · Cave of Flames · Fortress of Winds · Temple of Droplets · Royal Crypt · Palace of Winds · Dark Hyrule Castle|
|Smith's Sword · White Sword · Four Sword · Small Shield · Mirror Shield · Bombs · Boomerang · Bow · Cane of Pacci · Gust Jar|
|Bottle · Flame Lantern · Flippers · Grip Ring · Mole Mitts · Ocarina of Wind · Power Bracelets · Roc's Cape|
|Elements · Kinstones · Tiger Scrolls|
|Deku · Fairy · Goron · Human · Hylian · Minish · Wind Tribe|